Mammal Study

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Instructions to Authors

Mammal Study (Print ISSN: 1343-4152, Online ISSN: 1348-6160) is the international journal published quarterly by the Mammal Society of Japan. It is cited in Science Citation Index Expanded by Thomson Reuters, and publishes three types of contributions on all aspects of Mammalogy: Original paper, Short communication, and Review. Submission can be made by members and non-members of the society. Mammal Study especially encourages publications for the study of mammals in Asia. In addition to the printed journal, the electric version of Mammal Study can be accessed at BioOne ( Authors are recommended to follow Instructions to Authors and also refer to recent issues of the journal for details of style and layout.

Submission of manuscript

 Original papers, Short communications, and Reviews should be original and involve new scientific findings and ideas, and cannot have been published in or concurrently submitted to other journals or kinds of publications irrespective of written languages. A Short communication cannot exceed 3,500 words including references. All co-authors should read the submitted manuscript and agree on the submission. Corresponding author should be a single person and have been delegated the responsibility for the editorial process of the manuscript. Double corresponding authors are not permitted. All manuscripts that include sampling and handling animal specimens are required to follow the Guidelines for the Procedure of Obtaining Mammal Specimens as Approved by the Mammal Society of Japan ( Authors who are not native English speakers should seek English correction before submission. Supplementary materials online are not allowed to be used. There is no publication charge to the author except for color prints as mentioned below.
 Submitted manuscripts are subject to scientific review. Initial review is done by the Editor-in-Chief and by the associate editor in charge of the manuscript, who evaluate whether the manuscript is of sufficient quality and general interest for further review. Manuscripts that pass this initial evaluation will be sent to at least two qualified reviewers before the editor decides on publication. Authors can recommend preferred and non-preferred reviewers, but the reviewer selection is made by the associate editor in charge of the manuscript. Authors are notified within three months about the decision of acceptance, minor or major revision, encouraged resubmission, and reject, and provided with reviewers’ and associate editor’s comments. The revised manuscript is required to be re-submitted within 90 days after the notification.
 Authors should submit the manuscripts via ScholarOne Manuscripts online submission system for Mammal Study. The combined size for all uploaded files should be no larger than 10 MB. The URL is as follows.

 To submit your manuscript, first create an account on Scholar One Manuscripts, and then access the Author Center. Click on the ‘Submit New Manuscript’ and follow the instructions. All necessary information should be provided, and the transformed PDF file for review purposes should be carefully checked by the authors before finalizing the submission. Incompleteness of authors’ information or PDF transformation errors may cause delay of the review process.
All correspondences regarding manuscripts and editorial matters should be addressed to:
  Editor-in-Chief, Takuya Shimada, PhD
  Tohoku Research Center, Forestry and Forest Products
  Research Institute
  92-25 Nabeyashiki, Shimo-Kuriyagawa, Morioka, Iwate
  020-0123 Japan
  TEL: +81-19-641-2150
  E-mail: mammal_study[AT]

Proof and reprints

Galley proof and the reprint order form will be sent to the corresponding author. Reprints may be purchased in blocks of 50. PDF file of published paper is provided to the corresponding author free of charge. All the communication for proof and reprints will be made using e-mail between author, editor, and printing company.


 The copyrights for articles in Mammal Study are held by the Mammal Society of Japan. For the purpose of an institutional repository, authors can upload the Mammal Study version (not private copies) after 1 year of publication.

Form of manuscripts

 Manuscript text file should be prepared as Microsoft Word in the A4 size (21.0 × 29.7 cm). Margin should be 30 mm on all sides for all submitted pages. Do not hyphenate words at the right margin. Text should be 12 pt and line breaks should be 24 pt. Text pages should be numbered in the center of the footer. Line number should be given in the left margin for every fifth line (e.g. 1, 5, 10, 15, etc.). Each sentence should conclude with a full stop followed by a single space.
 Manuscript text file should be arranged in the following order: on the first page, the title, name(s) of author(s) and affiliation, running head (fewer than 40 letters, including spaces), total number of words including references, and e-mail address for correspondence; from the second page, abstract (fewer than 200 words) and key words (five words or fewer that are not found in the title), main text, acknowledgments, references, figure legends. Titles of papers must be accurate and concise, and (for abstract services) include any relevant taxonomic name. Tables and figures are not included in the manuscript file but will be submitted as separate files.


 Each table should be prepared separately by Microsoft Excel and uploaded individually.


 Each figure should be prepared separately and uploaded individually. Figures should be simple and self-explanatory. The recommended formats for figures are TIF, EPS, PSD (Adobe Photoshop) and AI (Adobe Illustrator). Color figures are charged JPY 43,200 for each page with color figures. Black and white (B/W) figures should be carefully prepared to be clear for the reader, and gray tone should be avoided as much as possible. If the authors choose to use color illustration, authors should select the color to increase clarity for the reader. For B/W figures, all the submitted files for review also should be B/W. Color photos and illustrations can be printed in B/W and published in color in the PDF for free, with the authorization of the associate editor in charge of the manuscript and editor-in-chief. In this case, figures should be informative even in B/W print. When authors request this option, both the color and B/W figures should be submitted. The resolution of color pictures should be a minimum of 300 dpi.

Measurements, abbreviations and others

 Authors are requested to use the International System of Units (SI, Système International d’Unités) where possible for all measurements (see SI Brochure: The International System of Units, Eighth edition, 2006, Arabic numerals should be used for numbers exceeding 10. Numerals of 10 or less than 10 should be spelled out. Scientific names, but not the taxonomic authority, should be written in italics.

 All details of statistical outcomes reported should be provided, and degrees of freedom should be reported as subscripts of test statistics (e.g., t2 = 3.76, P < 0.04; F6,198 = 0.253, P = 0.618).

Other statistical standards:
  SD, SE, df
    All tests in italics: F, G, H, P, R, r, R2, t, U, V, W, z, etc.
    Spell out chi-square test but use χ2 with values; e.g., we used a chi-square test, but χ2 = 234.55.
Define analysis of variance and other statistical acronyms (e.g., coefficient of variation, confidence interval, etc.) at first use; thereafter use ANOVA, CV, CI, etc.


References in the text, figure, table, and appendix
 References in the text, figure, table, and appendix should refer to: Single author: Motokawa (2008) / (Motokawa 2008)
 Two authors: Uchida and Shiraishi (1985) / (Uchida and Shiraishi 1985)
Three or more authors: Miura et al. (1993) / (Miura et al. 1993)
 In case where an exact verbatim quotation is provided, the authors can specify the page number, such as “Motokawa (2008: 73) stated that ...”.
 More than one reference within the same parentheses must be listed chronologically, and alphabetically if of the same year. In case that the authors cite both single author’s and his or her collaborative papers in the same year, the single author’s paper should be placed first. An example of references in parentheses in the text is: (Shimada2008; Motokawa et al. 2009; Oshida 2009; Oshida et al. 2009, 2010; Sato et al. 2010)
 The precise journal and page information must be provided for references cited in the Abstract according to the following example: Sato et al. (2014; Mammal Study 39: 1–10) / (Sato et al. 2014; Mammal Study 39: 1–10).

References in the References section

 All references cited in the text, figures, tables, and appendices must appear in the References section, and vice versa. References in this section should be listed alphabetically by the first author according to examples below. Unpublished manuscript (in preparation, submitted, or in review), unpublished data, and/or personal communication are not allowed to be listed in the References section, but may be mentioned in the text.
 If more than one reference from the same author(s) is cited, they must be distinguished by the letters a, b, c, etc. after the year (e.g., 1997a). If the first authors are the same and the other authorships are different among the papers of the same year, those references must be listed alphabetically according to the surname of the second or later authors. For papers with more than 10 authors, “et al” must be used after the 10th author name. The journal name must not be abbreviated. The issue number should be provided if the pagination is not consecutive between different issues. For references not written in English, (in Japanese), (in Japanese with English Abstract), or derivatives from these styles must be placed prior to the last period in the reference. References only for the authority for a taxonomic name should not be listed in the Reference section.

Examples for references

Journal Articles
Printed Journals:
Abe, H., Shiraishi, S. and Arai, S. 1991. A new mole from Uotsurijima, the Ryukyu Islands. Journal of the Mammalogical Society of Japan 15: 47–60.
Printed Journals in Online Early View:
Delibes-Mateos, M., Ferreira, C., Rouco, C., Villafuerte, R. and Barrio, I. C. 2014. Conservationists, hunters and farmers: the European rabbit Oryctolagus cuniculus management conflict in the Iberian Peninsula. Mammal Review. DOI: 10.1111/mam. 12022.
Online Journals with DOI:
Isaac, N. J. B., Turvey, S. T., Collen, B., Waterman, C. and Baillie, J. E. M. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PLOS ONE 2 (3): e296. DOI: 10.1371/journal.pone.0000296.
Online Journals without DOI:
Instead of providing DOI, the correct URL and the access date information such as (Accessed 30 May 2014) must be given.
Whole Books:
Eisenberg, J. F. 1981. The Mammalian Radiations. University of Chicago Press, Chicago, 610 pp.
Chapters in Books:
Shimada, T. 2015. Acorns as a resource for forest-dwelling mammals in Japan. In (Ohdachi, S. D., Ishibashi, Y., Iwasa, M. A., Fukui, D. and Saitoh, T., eds.) The Wild Mammals in Japan, Second edition, pp. 198–200. Shoukadoh, Kyoto.
Zwickl, D. J. 2006. Genetic Algorithm Approaches for the Phylogenetic Analysis of Large Biological Sequence Datasets under the Maximum Likelihood Criterion. Ph. D. Thesis, University of Texas at Austin, Austin, 114 pp.
Computer programs
If the program development was reported in a journal article, a book (or a book chapter), or a thesis, refer to examples described above. Otherwise, cite as:
Pagel, M. and Meade, A. 2007. BayesTraits. Available at (Accessed 19 January 2011).
R Development Core Team. 2011. R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, Vienna, Austria. Available at (Accessed 30 May 2014).
Online Sources
The International Union for Conservation of Nature. 2013. 2013 IUCN Red List of Threatened Species. Version 2013.1. Available at (Accessed 30 May 2014).
In Press Manuscript
 For a manuscript accepted but not yet published, “(in press)”must be given after the journal name.